There is always some water at the source of all the flooding. So even when the Okavango Delta (the famous delta in Africa I assume you’re talking about) mostly dries up, there are still places for fish to live up in the headlands. The source when the Okavango Delta floods is 1250 km away in the Angola highlands. Any fish which are present when the delta floods are only there because they came with the flood waters; they were previously living in rivers and lakes which overflowed with rainwater and spilled into the delta. Many species rely on the flood plains as a feeding and spawning ground. Check out the second citation for an in-depth discussion of the fish in the Okavango Delta.
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