Fish still breath oxegyn, they just get it from the water. If they are kept in a tank or even a vernal pool where the number of organisms living there depletes the oxegyn in the water, then the fish can in fact drown. That is one of the reasons that filters are used in aquariums. They keep the water flowing and oxegynated.
If there is no filter yes it can. Well yea basically everything neslam said
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