There’s a huge one up front: no light. Sunlight provides the energy source for photosynthetic organisms (plants, algae, etc). The raw energy of sunlight is the base for the vast majority of ecosystems on earth. Deep cave systems have no access to sunlight, so photosynthetic plants and algae don’t grow. These are the base for most terrestrial and aquatic ecosystems — certainly the most diverse ones. The organisms that live in caves need to find some way around this to get their energy. Bottom line, there’s just not much energy (i.e. food) available in caves.
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